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### Climate shifts the probability of developmental forms
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Figure 3 projects climate onto Fire VASE space. Event-mean VPD varies across the developmental axes, and composite VASEs show that low-, middle-, and high-VPD fires differ in where normalized growth is allocated through developmental time. Developmental-neighborhood prevalence shifts across VPD groups, and effect-size summaries show that maximum temperature, VPD, relative humidity, fuel moisture, precipitation, and fire-danger indices relate to different developmental responses. These associations are coherent with the broader literature linking warming, aridity, and fuel dryness to fire activity (1-3), but the VASE analysis resolves the outcome as a developmental distribution rather than a single aggregate burned-area response.
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Figure 3 projects climate onto Fire VASE space. Climate-colored morphospace maps show that maximum temperature, VPD, fuel moisture, and wind emphasize different portions of the developmental space, while composite VASEs show that low-, middle-, and high-VPD fires differ in where normalized growth is allocated through developmental time. Developmental-neighborhood prevalence shifts across VPD groups, and effect-size summaries show that maximum temperature, VPD, relative humidity, fuel moisture, precipitation, and fire-danger indices relate to different developmental responses. These associations are coherent with the broader literature linking warming, aridity, and fuel dryness to fire activity (1-3), but the VASE analysis resolves the outcome as a developmental distribution rather than a single aggregate burned-area response.
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The predictive limit is equally important. In conservative blocked validation, which summarizes transfer across year, region, and region-year blocks rather than random-fire splits alone, the best transferable event-level representation is core event means, with median held-out R2 of 0.349 across developmental responses. Region-season anomaly diagnostics do not outperform core event means, comprehensive event means do not outperform core event means, and temporally resolved exposure summaries do not outperform core event means in the median blocked comparison. This does not mean that humidity, fuel moisture, precipitation, or fire danger are unimportant. It means that, in these correlated daily centroid summaries and linear blocked baselines, adding more climate descriptors did not improve transfer beyond the core atmospheric variables. Modest blocked R2 is therefore a bound on deterministic prediction, not a rejection of the distributional result. The claim is probabilistic: climate redistributes fires across developmental possibilities. It does not assign a unique developmental form.
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